Seahorse::Client::NetworkingError: Empty or incomplete response body (Seahorse::Client::NetworkingError)

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When running the command sudo -E -u discourse bundle exec rake s3:upload_assets

I am using Cloudflare R2, configures as S3, from this post:

I originally tried the same command, sudo -E -u discourse bundle exec rake s3:upload_assets but it resulted in:

/root is not writable.
Bundler will use /tmp/bundler20250409-510203-w6snye510203 as your home directory temporarily.
ERROR: Ensure S3 is configured in config/discourse.conf or environment vars

So, I put the configuration in as environment variables in app.yml - which looks like:

## S3 Configuration
  DISCOURSE_USE_S3: true
  DISCOURSE_S3_REGION: us-east-1
  DISCOURSE_S3_ACCESS_KEY_ID: xxx
  DISCOURSE_S3_SECRET_ACCESS_KEY: xxx
  DISCOURSE_S3_CDN_URL: https://pub-xxx.r2.dev
  DISCOURSE_S3_ENDPOINT: https://xxx.r2.cloudflarestorage.com/xxx
  DISCOURSE_S3_BUCKET: xxx

It yields the following error:

/root` is not writable.
Bundler will use `/tmp/bundler20250409-1393-48ek5u1393' as your home directory temporarily.
Installing CORS rules...
Attempting to apply ASSETS S3 CORS ruleset in bucket biohacking-forum.
rake aborted!
Seahorse::Client::NetworkingError: Empty or incomplete response body (Seahorse::Client::NetworkingError)
/var/www/discourse/vendor/bundle/ruby/3.3.0/gems/aws-sdk-core-3.219.0/lib/seahorse/client/plugins/raise_response_errors.rb:17:in `call'
/var/www/discourse/vendor/bundle/ruby/3.3.0/gems/aws-sdk-s3-1.182.0/lib/aws-sdk-s3/plugins/sse_cpk.rb:24:in `call'
/var/www/discourse/vendor/bundle/ruby/3.3.0/gems/aws-sdk-s3-1.182.0/lib/aws-sdk-s3/plugins/dualstack.rb:21:in `call'
/var/www/discourse/vendor/bundle/ruby/3.3.0/gems/aws-sdk-s3-1.182.0/lib/aws-sdk-s3/plugins/accelerate.rb:43:in `call'
/var/www/discourse/vendor/bundle/ruby/3.3.0/gems/aws-sdk-core-3.219.0/lib/aws-sdk-core/plugins/checksum_algorithm.rb:169:in `call'
/var/www/discourse/vendor/bundle/ruby/3.3.0/gems/aws-sdk-core-3.219.0/lib/aws-sdk-core/plugins/jsonvalue_converter.rb:16:in `call'
/var/www/discourse/vendor/bundle/ruby/3.3.0/gems/aws-sdk-core-3.219.0/lib/aws-sdk-core/plugins/invocation_id.rb:16:in `call'
/var/www/discourse/vendor/bundle/ruby/3.3.0/gems/aws-sdk-core-3.219.0/lib/aws-sdk-core/plugins/idempotency_token.rb:19:in `call'
/var/www/discourse/vendor/bundle/ruby/3.3.0/gems/aws-sdk-core-3.219.0/lib/aws-sdk-core/plugins/param_converter.rb:26:in `call'
/var/www/discourse/vendor/bundle/ruby/3.3.0/gems/aws-sdk-core-3.219.0/lib/seahorse/client/plugins/request_callback.rb:89:in `call'
/var/www/discourse/vendor/bundle/ruby/3.3.0/gems/aws-sdk-core-3.219.0/lib/aws-sdk-core/plugins/response_paging.rb:12:in `call'
/var/www/discourse/vendor/bundle/ruby/3.3.0/gems/aws-sdk-core-3.219.0/lib/seahorse/client/plugins/response_target.rb:24:in `call'
/var/www/discourse/vendor/bundle/ruby/3.3.0/gems/aws-sdk-core-3.219.0/lib/aws-sdk-core/plugins/telemetry.rb:39:in `block in call'
/var/www/discourse/vendor/bundle/ruby/3.3.0/gems/aws-sdk-core-3.219.0/lib/aws-sdk-core/telemetry/no_op.rb:29:in `in_span'
/var/www/discourse/vendor/bundle/ruby/3.3.0/gems/aws-sdk-core-3.219.0/lib/aws-sdk-core/plugins/telemetry.rb:53:in `span_wrapper'
/var/www/discourse/vendor/bundle/ruby/3.3.0/gems/aws-sdk-core-3.219.0/lib/aws-sdk-core/plugins/telemetry.rb:39:in `call'
/var/www/discourse/vendor/bundle/ruby/3.3.0/gems/aws-sdk-core-3.219.0/lib/seahorse/client/request.rb:72:in `send_request'
/var/www/discourse/vendor/bundle/ruby/3.3.0/gems/aws-sdk-s3-1.182.0/lib/aws-sdk-s3/client.rb:6359:in `get_bucket_cors'
/var/www/discourse/lib/s3_helper.rb:401:in `fetch_bucket_cors_rules'
/var/www/discourse/lib/s3_helper.rb:168:in `ensure_cors!'
/var/www/discourse/lib/s3_cors_rulesets.rb:60:in `sync'
/var/www/discourse/lib/tasks/s3.rake:183:in `block in <main>'
/var/www/discourse/vendor/bundle/ruby/3.3.0/gems/rake-13.2.1/exe/rake:27:in `<top (required)>'
/usr/local/bin/bundle:25:in `load'
/usr/local/bin/bundle:25:in `<main>'
Tasks: TOP => s3:upload_assets => s3:ensure_cors_rules
(See full trace by running task with --trace)```