I don’t get this anymore in 2.7.3 but therefore now getting.
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Do you use more than one email address? Did you reply from a different email address? Email replies require that you use the same email address when replying. Alternately, the Message-ID header in the email may have been modified.
Seems to happen if other replies where made after the post that I am replying to.
Since this seems difficult to debug (user error, replying to From instead to Reply-To - no I did not do that in this case / mail server messing something up and/or discourse bug), I came up with the following feature request to ease debugging:
Reply with debug info to rejected e-mails